Three things stood out on the defense on Saturday:
1) Coples needs to improve at setting the edge against the run. He was caught out of position several times. Experience will help with that, and he even adjusted to it as the game went on. Unfortunately his injury will delay his progression.
2) Damon Harrison was awful. Completely failed to anchor the line, and didn't command double teams. That weakens the whole middle of the defense tremendously.
3) Milliner played poorly. He was so-so in coverage, but terrible against the run. On one run play in particular, he was unblocked, but allowed the runner to get outside along the sideline for a big gain. If he had forced the runner inside, he would have been tackled for no gain by one of the linebackers.
Milliner and Coples' issues were due to inexperience. They can be fixed. But Harrison just flat-out got his ass kicked.